The fate of "hidden" Stone of Destiny fragments has been traced by a researcher who has been unravelling the mysteries behind the ancient artefact.
Professor Sally Foster's theory about the puzzling "XXXV" Roman numeral on the underside of the sandstone block has also been published in an academic journal.
The Stirling University academic has been carrying out painstaking research to uncover the stories behind several fragments which were separated from the original piece when it was repaired in secret in 1951.
The 152kg block was spirited away from Westminster Abbey by a group of Scottish nationalist students the previous year.
It broke in two during the raid and was repaired by stonemason and sympathiser Bertie Gray before being returned to the police.
Prof Foster's research suggests Gray collected 34 fragments of the original stone during this secretive work, as well as potentially inscribing the numeral for 35 on the stone itself as a light-hearted nod to his own role in the story of the artefact.
Following a public appeal earlier this year, her research has been published in The Antiquaries Journal as an academic paper.
The lives of several fragments are traced, some of which were gifted to SNP politicians, including the late former first minister Alex Salmond and two former MPs.
One fragment travelled halfway across the world to Australia and is stored in a museum.
Prof Foster said: "This is not just any stone.
"When Scottish nationalists manhandled it out of the Coronation Chair and secreted it away from London's Westminster Abbey on Christmas morning 1950, this caused the English-Scottish border to be closed for the first time in 400 years, because, since the 14th century, nearly all English, later British monarchs, sat over the Stone during their coronation, in an act that symbolised the subjugation of the Scots."
The stone, also known as the Stone of Scone, currently sits as the centrepiece exhibition of Perth Museum, which narrates its centuries-long history with the Scottish and later British royal families.
The existence of fragments from the Stone of Destiny has already been acknowledged - an inch-sized piece was gifted to Mr Salmond by Neil MacCormick and was kept at SNP headquarters - but it was only Prof Foster's research which revealed there were more than 30 in existence.
The Stirling University researcher also discovered that Canadian journalist and Calgary Herald editor Dick Sanburn was gifted a fragment and had it mounted behind his desk.
Prof Foster said: "With the likely perception of the fragments as being stolen property, few people opted to brazenly flaunt and taunt with their possession, except for some politicians.
"Families cared for them, emotionally and physically, and we can also trace the progression of fragments to valued heirlooms."
When Britain's Prince Charles was coronated in 2023 the Stone of Destiny was taken to London where it was mounted in the Coronation Chair, continuing its ancient role in the monarchy.
It moved to Perth Museum last year, having previously been kept at Edinburgh Castle.
Describing the mysterious Roman numerals on the stone, Prof Foster said: "I believe Gray, having gathered the 34 fragments, applied the XXXV as part of his finishing touches to the Stone's repair - the fragments plus the Stone equals 35.
"He used Roman numerals because these are easier to scratch into stone and, as a mason, he would have been familiar with historic masons' marks.
"He had the rare and almost unique opportunity to access the underside of the Stone, he had the means, he had the sense of humour, and the number 35 would only have been meaningful and significant to him."
