Al-Qaeda in the Arabian Peninsula has said that a US air strike had killed the senior figure who issued the group's claim of responsibility for the Charlie Hebdo attack in Paris.
The SITE Intelligence Group quoted al Qaeda as saying in an online video that AQAP ideologue Nasser bin Ali al-Ansi was killed with his eldest son and other fighters in Yemen.
Ansi's reported death suggests the covert US drone programme against the Yemen branch of the global militant group is continuing.
This is despite the evacuation of American military advisers from the country amid a worsening civil war.
Ansi, an ideologue and former fighter, had appeared in several of the group's videos.
In a message on 14 January, he said of the 7 January attack in Paris that the "one who chose the target, laid the plan and financed the operation is the leadership of the organisation", without naming an individual.
A week later he called for lone-wolf attacks in Western countries like America, Britain, Canada and France, as such operations were "better and more harmful".
In the Paris attack, 17 people, including journalists and police, were killed in three days of violence, including a mass shooting at the weekly Charlie Hebdo, known for its satirical attacks on Islam and other religions.
The attackers, two French-born brothers of Algerian origin, singled out the magazine for its publication of cartoons depicting and ridiculing the Prophet Mohammed.
The bloodshed ended on 9 January with a hostage-taking at a Jewish shop in which four hostages and the gunman were killed.
Ansi had also called for Yemeni Sunnis to confront the Houthi militia which has taken over large parts of Yemen since September.
Al-Qaeda views Houthis as heretics since they belong to a branch of Shia Islam.
SITE cited media as saying Ansi was killed in a drone strike in Mukalla, a city in Yemen's Hadramawt governorate, in April,along with his son and six other fighters.
Ansi had fought in Bosnia in the 1990s and worked for al-Qaeda in the Philippines and Afghanistan.
On 14 April AQAP announced that one of its leaders, Ibrahim al-Rubaish, had been killed by a US air strike.
Rubaish was a Saudi national released from the Guantanamo Bay prison camp in 2006.
The United States has poured aid and personnel into Yemen in recent years as part of its war on Islamist militants.
But it withdrew military personnel and pulled out of the al-Anad military base it was using in Yemen last month, as Iran-allied Shi'ite Muslim Houthi fighters advanced, plunging the impoverished country further into chaos.