England have confirmed that Chris Robshaw will remain captain for next month's autumn internationals at Twickenham, but there is no place for Saracens' Chris Ashton of Sale's Danny Cipriani in the squad.
Robshaw has been under pressure for his place in the team but has been retained as skipper for the Tests against New Zealand, South Africa, Samoa and Australia.
Head coach Stuart Lancaster has included Bath wing Semesa Rokoduguni and Saracens lock George Kruis in his 33-man squad for the QBE Series. Rokoduguni has been promoted at the expense of Ashton.
"Chris Robshaw continues to lead from the front," Lancaster said. "With other players now stepping up into captaincy and senior roles at their clubs, we are building a team of leaders who can take England forward."
Manu Tuilagi has been omitted from the 33-strong party, but tellingly he has also been left out of the names not considered for squad selection due to injury.
Tuilagi was on Tuesday ruled out of all four of next month's Tests by his club Leicester because of a groin problem, but it is clear England hope he can yet play a role.
There is also no place for Cipriani, who had been touted to make a remarkable return the international wilderness.
"We have included Jonathan Joseph as an injury replacement for Manu," Lancaster said.
"It's obviously disappointing to lose Manu but Jonathan has been in great form and deserves this chance.
"Our medics will continue to work closely with Leicester's and ensure that Manu is supported in his rehabilitation from his injury."
England senior squad (33) for forthcoming autumn internationals:
Forwards: J Marler (Harlequins), M Mullan (Wasps), K Brookes (Newcastle), D Wilson (Bath), D Hartley (Northampton), R Webber (Bath), D Attwood (Bath), G Kruis (Saracens), J Launchbury (Wasps), C Lawes (Northampton), C Clark (Northampton), J Haskell (Wasps), B Morgan (Gloucester), C Robshaw (Harlequins, capt), B Vunipola (Saracens), T Wood (Northampton).
Backs: D Care (Harlequins), L Dickson (Northampton), B Youngs (Leicester), O Farrell (Saracens), G Ford (Bath), S Myler (Northampton), B Barritt (Saracens), L Burrell (Northampton), K Eastmond (Bath), J Joseph (Bath), B Twelvetrees (Gloucester), M Brown (Harlequins), A Goode (Saracens), J May (Gloucester), J Nowell (Exeter), S Rokoduguni (Bath), M Yarde (Harlequins).
