Mohaather demonstrated a devastating turn of foot to win a stellar renewal of the Qatar Sussex Stakes at Goodwood.
Trained by Marcus Tregoning and ridden by Jim Crowley, the four-year-old endured a luckless run in the Queen Anne Stakes at Royal Ascot - but righted the record as an impressive victor in a top-class running of the Group One showpiece.
Queen Anne winner Circus Maximus set out to make all under Ryan Moore and the field was closely bunched with two furlongs to run as the leader tried to press on, with San Donato the first to make his bid and briefly looking as though he could be set to produce a shock.
However, Ger Lyons' previously-unbeaten Siskin then unleashed his challenge down the outside, with Crowley producing his mount even wider still and once he did Mohaather (3-1) fairly flew in the final furlong to collar Circus Maximus and score a first top-level victory.
Irish 2,000 Guineas hero Siskin dropped back to third, with Kameko - winner of the Newmarket Guineas - fourth having been tight for room in the closing stages.
Crowley said: "The plan was to ride him a bit more forward today, we were aware it was going to be tactical. When I jumped out, I would have liked to have sat third or fourth, one off the rail and kept my options open.
"The pace was very slow and Frankie (Dettori, on Wichita) arrived on my outside because we were going so slow and we were in a bit of a pocket and I knew then we were going to need a bit of luck in running.
"We had to switch around and he just showed the most electric turn of foot. To give weight away to younger horses and pick them up the way he did was special really.
"I knew I probably wasn't going to get a run as they'd all rolled back in towards the rail. I could see Siskin on the outside and I knew he was going to play his cards late, so my option then was to try to get on the back of him and follow him through."
Looking back on Mohaather's defeat in the Queen Anne, Crowley said: "I was gutted, it was eating away at me for a few days afterwards. When he came out again at Ascot (in the Summer Mile), I was so happy to see the way he won that day because I'd ridden him work a few times early on and he'd given me a feel like not many horses have done.
"He's so athletic, he's a beautiful-looking horse, he's got class and we've always thought he's got so much pace that we even contemplated early on going sprinting with him.
"He'd win over six, but today shows it was the right decision to go down the mile route and that's the sign of a really good horse - he's got so much pace."
Steel Bull repaid Curragh trainer Michael O'Callaghan's faith and adventure with victory in the Markel Insurance Molecomb Stakes.
Well-backed as the 2-1 favourite following his Naas maiden victory just a week ago, the Colin Keane-ridden Clodovil colt had to wait behind horses for quite some time but arrived on the scene approaching the final furlong and quickly moved past the pace-setting Ben Macdui.
He was then not hard-pressed to hold on to his advantage, giving O'Callaghan his highest-profile British success to date in this Group Three sprint for juveniles.
Ben Macdui stayed on to finish a 16-1 runner-up, with outsider Internationaldream taking third a length and a half further back.
Keane said: "Looking at the replay of his maiden, he won it very well - and talking to Michael beforehand, he was very confident that he had a lot of speed.
"He showed a very good turn of foot today. He puts his head down and tries very hard, and he's a horse that is going to keep progressing.
"I'd imagine he would get an extra furlong, because he settles. I suppose the track today was an unknown, but he handled it very well."
O'Callaghan said: "He did it very well, and we're delighted with him.
"We knew he was a very nice colt. He didn't get a clear passage today, but he quickened up like a smart one and got a very good ride.
"He only won his maiden a week ago, so there should be more improvement in him, and I think he'd stay six furlongs as he settles so well."
Steel Bull currently races in the trainer's colours, but O'Callaghan admits he is open to selling the youngster if the price is right.
He said: "We'll see what happens. There was a bit of interest before today, but not enough to prise him away for us.
"Hopefully that might change after today, but either way we're very happy to have him."
Earlier, Mambo Nights provided jockey Andrea Atzeni and owner Sheikh Mohammed Obaid with a timely boost on a difficult day by continuing his upward curve in the Unibet 3 Boosts A Day Handicap at Goodwood.
The Richard Hannon-trained Havana Gold colt made it a hat-trick of wins for the season when holding A Star Above at bay by a neck in the mile-and-a-half prize.
Victory for Mambo Nights came just hours after news of the death of the high-profile owner's multiple Group-race winner Defoe, who suffered a fatal injury on the gallops in Newmarket.
Atzeni said of the 9-1 winner: "He is a big, raw horse. He won quite nicely at Ripon the other day and he had quite good two-year-old form.
"He is a work in progress and Richard has done a great job with him, taking it step by step. He will be even better at four, I'd say."
Paying tribute to Defoe, he added: "It is very sad news losing any horse really, but Defoe was very close to us. Sheikh Mohammed Obaid bred the horse so knew him as a foal.
"I remember like it was yesterday flying to Ffos Las to ride him first time out when he won his maiden. He is a horse that gave us a lot of great days. He won the Coronation Cup, won the Hardwicke and ran well in Dubai.
"I was hoping one day when he retired I could have him in my paddocks so I could look after him in his older days, but unfortunately it wasn't meant to be."
Just Herbert demonstrated stamina is his forte when leaving his previous efforts this season behind to spring a surprise in the Unibet You're On Handicap.
After failing to beat a rival on his return at Haydock, the William Muir-trained four-year-old had continued his recent spell in the doldrums when finishing down the field in both the Northumberland Plate at Newcastle and last time out in the Marsh Cup at Newbury.
Looking a completely different character to the one seen in his previous three starts, the 25-1 Dunaden gelding appeared to appreciate every yard of the extended two-and-a-half-mile trip to wear down long-time leader Rochester House almost on the line before scoring by a neck.
Ralph Beckett could pitch Chamade (22-1) back into Listed company on her next start after the daughter of Sepoy opened her account for the campaign with a front-running success in the British Stallion Studs EBF Fillies' Handicap.