The car is a white Audi A6 'Allroad' 3.0 TDI; it rolled off the assembly line at the Neckarsulm plant.
The record figure is a testament to the leading role played by the pioneering all-wheel drive.
No other premium brand in the world can claim such high sales figures for a four-wheel drive or a comparable range of models boasting more than 140 Quattro variants.
Quattro technology debuted in 1980 in the legendary, original Quattro car.
Adding together all of the available body types, engines and drive systems amounts to more than 140 different models.
Audi offers the widest range of cars by far with all-wheel drive: the Audi Q7, R8, A4 Allroad and A6 Allroad, as well as all S and RS models, are available exclusively with Quattro drives - Audi offers it as an option in all other model lines.
Mark O'Connell, General Manager of Sales of Audi Ireland, said: "Technology and innovation is at the heart of the Audi ethos and it is a great achievement to have reached this milestone today of five million Audi Quattro drive systems sold."